Mactans is one of the Ancient Spider Queens and one of the main villains in Kingdom Rush: Origins.
Mactans first appears in stage 11, The Unseelie Court. After Malicia is defeated and incapacitated, Mactans suddenly descends from the sky, wrap Malicia in a web, and bring her away to the elven First City.
Mactans is later encountered in Mactans' Retreat. At several points, she'll descend onto the map, webbing towers. She can be tapped on to stop her short of disabling towers.
At the Shrine of Elynie, Mactans is seen working with Malicia to break the shield around the Tear of Elynie, the priceless artefact that gave birth to the elves. Much like in the previous level, she will web random towers, but she can also land on a path, covering it with more web, laying Spiderbrood eggs (luckily, the Spiderbroods that spawn do grant gold when killed), and webbing any nearby soldiers (identical to the ability of Webspitter Spiders), and will also taunt you, along with Malicia. As before, she can be tapped to prevent her from disabling towers, but she will not fail to cover the path with web, deposit eggs, or web soldiers regardless of how quickly she is tapped.
After wave 15, the shield around the Tear of Elynie is broken. Both Malicia and Mactans jump at the Tear screaming 'Mine!', causing the Tear to become corrupted and fuse them into the Spider Goddess, the final boss of the campaign.
- (level start) It is good you made it this far...
- I excel at pulling strings!
- Strand by strand a web is spun!
- If only Sarelgaz could see this!
- I'm every nightmare you've ever had!
- Was that a twitch on my web?
- I'm your worst dream come true!
- Don't think too much! You'll die either way!
- Hizz...This is taking too long!
- I'm everything you ever were afraid of!
- Why didn't we do this earlier?